Okay! Here are the roles:

The Mudwing replacement, Angleworm
[TAKEN BY ME]

The Sandwing replacement, Sage
[WingsOfFire15]

The Seawing replacement, Lunople
[Aspen_The_Random_One]

The Nightwing replacement, Quickdeath
[GamethingZ20]

The Icewing replacement, Mistral
[ANRawesome ]

The Skywing replacement, Misty
[dollbaby_102]

-Guardian Roles-
THE SANDWING GUARDIAN: Sahara. (Teaches science, and math.)

THE NIGHTWING GUARDIAN, Duskbringer: [WingsOfFire15 ]. (Teaches music and art.)

THE SKYWING GUARDIAN: OPEN. (Teaches history, and battle training.)

THE ICEWING "GUARDIAN": [TAKEN BY ME]

-Other Roles-
THE WINGS OF NIGHT: TAKEN - me.

THE WINGS OF NIGHT'S MOST LOYAL FOLLOWER: OPEN (preferably a Nightwing or Night/hybrid.)

THE WINGS OF NIGHT'S SPY/ASSASSIN: TAKEN [TimeTrxveller] (any tribe)

THE WINGS OF NIGHT'S CHAMPION: OPEN (the most likely to die for Wings of Night and the rest of the Night army. A good fighter. Any tribe.)

THE WINGS OF SKY: OPEN

THE WINGS OF ICE: OPEN

THE WINGS OF SEA: OPEN

THE WINGS OF MUD: OPEN

THE WINGS OF SAND: OPEN

Random characters:
Cinder [WingsOfFire15 ]
Also you could just make a side character that chills in a kingdom if ya want-

Might make more roles, depending on who's interested :)