25. Icewings,(Skywings?) and Sandwings might have been the original animus users, let me explain.

Any Nightwing Animus would be related to Darkstalker, so Stonemover and Sunny. Darkstalker got his power from his Icewing father, so Stonemover and Sunny are part Icewing.

Albotross the Seawing is the first known Seawing and it was confirmed that he has Icewing blood, so Prince Fathom, Turtle and Princess Anemone would have a speck of Icewing blood though Albotross. 

And the Sandwing Animuss are Jeraboa 1 & 2. Though I can't explain the Skywing animus's